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thought. Teleology had become a bugbear to the vast majority of biologists, and all real understanding of the Cuvierian attitude seems, in most cases, to have been lost, although, curiously enough, teleological conceptions were often unconsciously introduced in the course of discussions on the "utility" of organs in the struggle for existence. Evolutionary morphology, being for the most part a form of pure or non-functional morphology, agreed then in all essential respects with pre-evolutionary or transcendental morphology. But it contained the germ of a new conception which threw a new light upon the whole science of morphology. This was the conception of the organism as an historical being. We have seen this thought expressed with the utmost clearness by Darwin himself (_supra_, p. 233). In his eyes the structure and activities of the living thing were a heritage from a remote past, the organism was a living record of the achievements of its whole ancestral line. What a light this conception threw upon all biology! "When we no longer look at an organic being as a savage looks at a ship as something wholly beyond his comprehension; when we regard every production of Nature as one which has had a long history; when we contemplate every complex structure and instinct as the summing-up of many contrivances, each useful to the possessor, in the same way as any great mechanical invention is the summing-up of the labour, the experience, the reason, and even the blunders of numerous workmen; when we thus view each organic being, how far more interesting--I speak from experience--does the study of natural history become!" (_Origin_, 6th ed., pp. 665-6). Sedgwick expressed the same thing from the morphological point of view when he wrote, with reference to the ancestral significance of the blastopore:--"If there is anything in the theory of evolution, every change in the embryo must have had a counterpart in the history of the race, and it is our business as morphologists to find it out" (p. 49, 1884). By the evolution-theory the problems of form were linked indissolubly with the problem of heredity. Unity of plan could no longer be explained idealistically as the manifestation of Divine archetypal ideas; it had a real historical basis, and was due to inheritance from a common ancestor. The evolution-theory gave meaning and intelligibility to the transcendental conception of the unity of plan; in particular it suppli
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