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divisions of the animal and vegetable kingdoms are much less close than the theory [of evolution] demands. Even the Archaeopteryx, the discovery of which made so much stir and appeared to establish a genetic relation between classes so distinct as Birds and Reptiles, fills up the gap only imperfectly, and does not indicate the point of bifurcation of these two classes. Intermediate links are lacking between Amphibia and Reptiles. Mammals, too, occupy an isolated position, and no zoologist can deny that they are clearly demarcated from other Vertebrates; indeed, no fossil mammal is certainly known which comes nearer to the lower Vertebrates than does Ornithorhynchus at the present day" (p. 115). To take a parallel from the Invertebrata, B. B. Woodward,[545] after discussing the phylogeny of the Mollusca as worked out by the morphologists and comparing it with the probable actual course of the evolution of the group, as evidenced by fossil shells, sums up as follows:--"The lacunae in our knowledge of the interrelationships of the members of the various families and orders of Mollusca are slight however, compared with the blank caused by the total absence from palaeontological history of any hint of passage forms between the classes themselves, or between the Mollusca and their nearest allies. Nor is this hiatus confined to the Molluscan phylum; it is the same for all branches of the animal kingdom. There is circumstantial evidence that transitional forms must have existed, but of actual proof none whatever. All the classes of Mollusca appear fully fledged, as it were. No form has as yet been discovered of which it could be said that it in any way approached the hypothecated prorhipidoglossate mollusc, still less one linking all the classes" (p. 79). Pointing in the same direction as the absence of transitional forms is the undeniable fact that all the great groups of animals appear with all their typical characters at a very early geological epoch. Thus, in the Silurian age a very rich fauna has already developed, and representatives are found of all the main Invertebrate groups--sponges, corals, hydroid colonies, five types of Echinoderms, Bryozoa, Brachiopods, Worms, many types of Mollusca and Arthropoda. Of Vertebrates, at least two types of fish are present--Ganoids and Elasmobranchs. In the very earliest fossiliferous rocks of all, the Precambrian formation, there are remains of Molluscs, Trilobites and Gigantost
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