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cases of this kind is recorded by Wigand in the 'Flora' for 1856, in a compound flower of _Polygonatum anceps_, in which within a twelve-parted perianth there were twelve stamens and two pistils, one four-celled, the other two-celled; hence it would appear as if a carpel belonging to one flower had become united to those constituting the pistil of the adjacent one. Among Orchids this fusion of some of the elements of different flowers, together with the suppression of others, is carried to such an extent as to render the real structure difficult to decipher. Sometimes flowers of _Ophrys aranifera_, at first sight seeming normal as to the number, and almost so as regards the arrangement of their parts, have yet, on examination, proved to be the result of a confluence of two flowers. Mr. Moggridge has observed similar phenomena in the same species at Mentone. Sometimes the fusion affects flowers belonging to different branches of the same inflorescence, as in _Centranthus ruber_, described by Buchenau, 'Flora,' 1857, p. 293, and even a blossom of one generation of axes may be united with a flower belonging to another generation. Thus M. Michalet[45] speaks of a case wherein the terminal flower of _Betonica alopecuros_ was affected with Peloria, and fused with an adjacent one belonging to a secondary axis of inflorescence, and not yet expanded. This latter flower had no calyx, but in its place were three bracts, surrounding the corolla; this again was united to the calyx of the terminal bloom in a most singular manner, the limb of the corolla and that of the calyx being so joined one to the other as to form but a single tube. It is not uncommon, as has been before stated, to find two corollas enclosed within one calyx, but this is probably the only recorded instance of the fusion of the calyx and corolla of two different flowers belonging to two different axes. From the preceding details, as well as from others which it is not necessary to give in this place, it would appear that synanthy is more liable to occur where the flowers are naturally crowded together[46] than where they are remote; so too, the upper or younger portions of the inflorescence are those most subject to this change. In like manner the derangements consequent on the coalescence of flowers are often more grave in the central organs, which are most exposed to pressure, and have the least opportunities of resisting the effects of that agency, than they
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