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in distinction to that of functional adaptation which interested Cuvier so greatly, came vividly before him as he worked through the vast collections of the museum. He was the first systematist to occupy himself in a philosophical manner with the problems of general biology. He introduced new problems and a new way of looking at old. With Lamarck the problem of species and the problem of ecological adaptation enter into general biology. The one point in which he does definitely carry on the thought of his predecessors is his conception of the animal kingdom as forming a scale of (functional) perfection. He did not go to the same extreme as Bonnet; he did not even consider that the animal series was a continuation of the vegetable series; in his opinion they formed two diverging scales. He recognised, too, that among animals there was no simple and regular gradation from the lowest to the highest, but that the orderly progression was disturbed and diverted by the necessity of adaptation to different environments. It is interesting to note that in developing this idea he arrived at a roughly accurate distinction between homologous and analogous structures. More importance, he thought, was to be attributed in classifying animals to characters which appeared due to the "plan of Nature" than to such as were produced by an external modifying cause (p. 299). But he did not formulate the distinction in any strictly morphological way. As his ideas developed he laid less stress upon the simplicity and continuity of the scale; in his supplementary remarks to the Introduction of 1816 he admits that the series is really very much branched, and even that there may be two distinct series among animals instead of one. His last schema of the course of evolution shows no little analogy with the genealogical trees of Darwinian speculation. It is headed "The presumed _Order_ of the formation of Animals, showing two separate partly-branching series," and it reads as follows:-- I.--_Series of Non-articulated_ II.--_Series of Articulated_ _Animals_. _Animals_. " I |-- Infusoria. n | | s A | Polyps. e n | | n i | ---------------- s m | | | i a | Ascidians. Radiates. Worms. t l | | | i s | | -
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