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thorough appreciation of the older classical or Cuvierian morphology that characterise Gegenbaur's work. According to Haeckel,[381] Gegenbaur was greatly influenced by J. Mueller, who, as we know, laid as much stress on function as on form. The "General Part" of Gegenbaur's text-book is in many ways a significant document and deserves close attention. We note first of all that physiology and morphology are considered by Gegenbaur to be entirely distinct sciences, with different subject-matter and different methods. "The task of physiology is the investigation of the functions of the animal body or of its parts, the referring back of these functions to elementary processes and their explanation by general laws. The investigation of the material substratum of these functions, of the form of the body and its parts, and the explanation of this form, constitute the task of Morphology" (2nd ed., p. 3). Morphology falls naturally into two divisions--comparative anatomy and embryology. The method of comparative anatomy is _comparison_ (p. 6), and in employing this method account is to be taken of "the spatial relations of the parts to one another, their number, extent, structure, and texture." Through comparison one is enabled to arrange organs in continuous series, and it comes out very clearly during this proceeding "that the physiological value of an organ is by no means constant throughout the different form-states of the organ, that an organ, through the mere modification of its anatomical relations, can subserve very different functions. Exclusive regard for their physiological functions would place morphologically related organs in different categories. From this it follows that in comparative anatomy we should never in the first place consider the function of an organ. The physiological value comes only in the second place into consideration, when we have to reconstruct the relations to the organism as a whole of the modification which an organ has undergone as compared with another state of it. In this way comparative anatomy shows us how to arrange organs in series; within these series we meet with variations which sometimes are insignificant and sometimes greater in extent; they affect the extent, number, shape, and texture of the parts of an organ, and can even, though only in a slight degree, lead to alterations of position" (p. 6). Geoffroy St Hilaire would have subscribed to every word of this vindicatio
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