thers with marbled
flowers. Gallesio[920] crossed reciprocally white and red carnations,
and the seedlings were striped; but some of the striped plants also
bore entirely white and entirely red flowers. Some of these plants
produced one year red flowers alone, and in the following year striped
flowers; or conversely, some plants, after having borne for two or
three years striped flowers, would revert and bear exclusively red
flowers. It may be worth mentioning that I fertilised the _Purple
Sweet-pea_ (_Lathyrus odoratus_) with pollen from the light-coloured
_Painted Lady_: seedlings raised from one and the same pod were not
intermediate in character, but perfectly resembled both parents. Later
in the summer, the plants which had at first borne flowers identical
with those of the _Painted Lady_, produced flowers streaked and
blotched with purple; showing in these darker marks a tendency to
reversion to the mother-variety. Andrew Knight[921] fertilised two
white grapes with pollen of the Aleppo grape, which is darkly
variegated both in its leaves and fruit. The result was that the young
seedlings were not at first variegated, but all became variegated
during the succeeding summer; besides this, many produced on the same
plant bunches of grapes which were all black, or all white, or
lead-coloured striped with white, or white dotted with minute black
stripes; and grapes of all these shades could frequently be found on
the same footstalk.

In most of these cases of crossed varieties, and in some of the cases of
crossed species, the colours proper to both parents appeared in the
seedlings, as soon as they first flowered, in the form of stripes or larger
segments, or as whole flowers or fruit of two kinds borne on the same
plant; and in this case the appearance of the two colours cannot strictly
be said to be due to reversion, but to some incapacity of fusion, leading
to their {394} segregation. When, however, the later flowers or fruit,
produced during the same season or during a succeeding year or generation,
become striped or half-in-half, &c., the segregation of the two colours is
strictly a case of reversion by bud-variation. In a future chapter I shall
show that, with animals of crossed parentage, the same individual has been
known to change its character during growth, and to revert to one of its
parents which it did not at first rese