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phology lies so far out of his ken that he has not been able to form an independent judgment of its aims and methods; but when, in spite of all this, he on every occasion lets fall a disparaging judgment of it, we must dispute his competence. It is true that in his Munich address he emphasises the statement, "That which graces me best is that I know my ignorance," by printing it in italics. I only regret that I am forced to deny his possession of this very grace. Virchow does not know how ignorant he is of morphology, else he would never have uttered his annihilating verdict on it, else he would not continually designate the study of the theory of descent as dilettanteism and vain dreaming, as "a fanciful private speculation which is now making its way in several departments of natural science." In truth, Virchow does me greatly too much honour when he designates as my "personal crotchet" an idea which for the last ten years has been the most precious common possession of all morphological science. If Virchow were not so unfamiliar with the literature of morphology, he must have known that it is penetrated throughout by this principle of descent, that every morphological inquiry which conscientiously pursues a well-considered problem now assumes the doctrine of descent as granted and indisputable. Of all this he is ignorant, and so it is intelligible that he should continue to demand "certain proofs" of this hypothesis, although those proofs have long since been produced. FOOTNOTES: [12] Vol. ii., p. 334 of translation. [13] London: C. Kegan Paul & Co. 1879. [14] Jena, Zeitschriften fuer Naturwissenschaft, 1875. Vol. x. Supplement. CHAPTER III. THE SKULL THEORY AND THE APE THEORY. Inasmuch as Virchow persists in treating the theory of descent as an "unproved hypothesis," inasmuch as he ignores all the forcible evidences of that hypothesis, he deprives himself of the right of speaking a decisive word in this, the most important scientific dispute of the present day. Virchow is, in fact, simply incompetent in the great question of evolution, as he is deficient in the greater part of that knowledge--more especially morphological knowledge--which is indispensable to forming a judgment upon it. Hence on the turning-point of the whole matter--viz., the problem as to the origin of species--he can have no opinion, as he has never turned his attention to the systematic treatment of species: those trans
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