change of pistils into stamens is more common in
monocarpellary pistils than it is in those which are made up of several
carpels. It seems clear that in this transformation the lobes of the
anther and the development of pollen have no relation to the production
of ovules.
=Staminody of the accessory organs of the flower.=--The scales that are
met with in some plants, either as excrescences from the petals, or as
imperfect representatives of stamens or other organs, are occasionally
staminoid; thus the scales of _Saponaria officinalis_, of _Silene_,
_Nerium Oleander_, the rays of _Passiflora_, the corona of _Narcissus_,
have all been observed occasionally to bear anthers.[333] In the case of
_Narcissus_ the loose spongy tissue of the corona seems to have the
nearest analogy to the anther-lobes, while the prolonged connective is
more like the ordinary segments of the perianth in texture. The species
in which this change may most frequently be observed are, _N. poeticus_,
_N. incomparabilis_, and _N. montanus_.
M. Bureau found in some flowers of _Antirrhinum majus_ two petal-like
bodies standing up in front of, or opposite to the two petals of the
upper lip,[334] and similar developments in which each of the two
adventitious segments are surmounted by an anther may be met with
frequently. It does not follow because these organs bear anthers that
they are morphologically true stamens. They are really scales, &c.,
taking on themselves accidentally the characters proper to stamens.
=Pistillody of the perianth.=--The passage of the segments of the
perianth into carpels has been observed frequently in _Tulipa
Gesneriana_, the change in question being generally attended by a
partial virescence. M. Gay is said by Moquin to have observed a flower
of _Crocus nudiflorus_ in which the segments of the perianth were cleft
and fringed at the same time, so that they presented the appearance of
the stigmas.
[Illustration: FIG. 160.--Flower of tulip, allowing vertical attachment
of a leaf, and also the existence of ovules on the margins of the
segments of the perianth. Some of the parts are removed.]
=Pistillody of the sepals.=--In some double flowers of the garden pea
communicated by Mr. Laxton, among other peculiarities was a
supernumerary 5-6-leaved calyx, some of the segments of which were of a
carpellary nature, and bore imperfect ovules on their margins, while at
their extremities they were drawn out into styles.[335]
=
|