ll be the tendency of
evolution in all organic systems subjected to restraints as those
we are acquainted with invariably are.

Such economy might be simply illustrated by considering the case
of a reservoir of water elevated above two hydraulic motors, so
that the elevated mass of water possessed gravitational
potential. The available energy here represents the stored-up
energy in the organism. How best may the water be conveyed to the
two motors [the organic systems reacting towards conditions A and
B] so

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that as little energy as possible is lost in transit? If the
motors are near together it is most economical to use the one
conduit, which will distribute the requisite supply of water to
both. If the motors are located far asunder it will be most
economical to lay separate conduits. There is greatest economy in
meeting a plurality of functions by the same train of
physiological processes where this is consistent with meeting
other demands necessitated by external or internal conditions.

But an important and obvious consequence arises in the supply of
the two motors from the one conduit. We cannot work one motor
without working the other. If we open a valve in the conduit both
motors start into motion and begin consuming the energy stored in
the tank. And although they may both under one set of conditions
be doing useful and necessary work, in some other set of
conditions it may be needless for both to be driven.

This last fact is an illustration of a consideration which must
enter into the phenomenon which an eminent biologist speaks of as
physiological or unconscious "memory,"[1] For the development of
the organism from the ovum is but the starting of a train of
interdependent events of a complexity depending upon the
experience of the past.

[1] Ewald Hering, quoted by Ray Lankaster, _The Advancement of
Science_, p. 283.

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In short, we may suppose the entire development of the plant,
towards meeting certain groups of external conditions,
physiologically knit together according as Nature tends to
associate certain groups of conditions. Thus, in the case in
point, climatic rigour and scarcity of pollinating agency will
ever be associated; and in the long experience of the past the
most economical physiological attitude towards both is, we may
suppose, adopted; so that the presence of one condition excites
the apparent unconscious memory of the other. In reality the
process of meeting the on