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108 experiment is not to be found in direct climatic influence upon the colour, but in causes which lie deeper, and involve some factors deducible from biological theory. The organism, as a result of the great facts of heredity and of the survival of the fittest, is necessarily a system which gathers experience with successive generations; and the principal lesson ever being impressed upon it by external events is economy. Its success depends upon the use it makes of its opportunities for the reception of energy and the economy attained in disposing of what is gained. With regard to using the passing opportunity the entire seasonal development of life is a manifestation of this attitude, and the fleetness, agility, etc., of higher organisms are developments in this direction. The higher vegetable organism is not locomotory, save in the transferences of pollen and seed, for its food comes to it, and the necessary relative motion between food and organism is preserved in the quick motion of radiated energy from the sun and the slower motion of the winds on the surface of the earth. But, even so, the vegetable organism must stand ever ready and waiting for its supplies. Its molecular parts must be ready to seize the prey offered to it, somewhat as the waiting spider the fly. Hence, the plant stands ready; and every cloud with moving shadow crossing the fields handicaps the shaded to the benefit of the unshaded plant in the adjoining field. The open bloom 109 is a manifestation of the generally expectant attitude of the plant, but in relation to reproduction. As regards economy, any principle of maximum economy, where many functions have to be fulfilled, will, we may very safely predict, involve as far as possible mutual helpfulness in the processes going on. Thus the process of the development towards meeting any particular external conditions, A, suppose, will, if possible, tend to forward the development towards meeting conditions B; so that, in short, where circumstances of morphology and physiology are favourable, the ideally economical system will be attained when in place of two separate processes, a, ss, the one process y, cheaper than a + ss, suffices to advance development simultaneously in both the directions A and B. The economy is as obvious as that involved in "killing two birds with the one stone"--if so crude a simile is permissible--and it is to be expected that to foster such economy wi
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