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study of domestic organisms to that of animals and plants in a state of nature. This is followed in both by a discussion of the _Difficulties on Theory_ and this by a section _Instinct_ which in both cases is treated as a special case of difficulty. If I had to divide the _Origin_ (first edition) into two parts without any knowledge of earlier MS., I should, I think, make Part II begin with Ch. VI, _Difficulties on Theory_. A possible reason why this part of the argument is given in Part I of the Essay of 1842 may be found in the Essay of 1844, where it is clear that the chapter on instinct is placed in Part I because the author thought it of importance to show that heredity and variation occur in mental attributes. The whole question is perhaps an instance of the sort of difficulty which made the author give up the division of his argument into two Parts when he wrote the _Origin_. As matters stand Sec.Sec. IV. and V. of the 1842 Essay correspond to the geological chapters, IX and X, in the _Origin_. From this point onwards the material is grouped in the same order in both works: geographical distribution; affinities and classification; unity of type and morphology; abortive or rudimentary organs; recapitulation and conclusion. In enlarging the Essay of 1842 into that of 1844, the author retained the sections of the sketch as chapters in the completer presentment. It follows that what has been said of the relation of the earlier Essay to the _Origin_ is generally true of the 1844 Essay. In the latter, however, the geological discussion is, clearly instead of obscurely, divided into two chapters, which correspond roughly with Chapters IX and X of the _Origin_. But part of the contents of Chapter X (_Origin_) occurs in Chapter VI (1844) on Geographical Distribution. The treatment of distribution is particularly full and interesting in the 1844 Essay, but the arrangement of the material, especially the introduction of Sec. III. p. 183, leads to some repetition which is avoided in the _Origin_. It should be noted that Hybridism, which has a separate chapter (VIII) in the _Origin_, is treated in Chapter II of the Essay. Finally that Chapter XIII (_Origin_) corresponds to Chapters VII, VIII and IX of the work of 1844. The fact that in 1842, seventeen years before the publication of the _Origin_, my father should have been able to write out so full an outline of his future work, is very remarkable. In his Autobiography{2
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