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tes, induces vegetative growth. This can be effected by culture in feeble light or in light deprived of the yellow-red rays: on the other hand, flower-production follows an increase in light-intensity. These results are essentially in agreement with well-known observations on cultivated plants, according to which, the application of much moisture, after a plentiful supply of manure composed of inorganic salts, hinders the flower-production of many vegetables, while a decrease in the supply of water and salts favours flowering. ii. INFLUENCE OF THE ENVIRONMENT ON THE FORM OF SINGLE ORGANS. (A considerable number of observations bearing on this question are given by Goebel in his "Experimentelle Morphologie der Pflanzen", Leipzig, 1908. It is not possible to deal here with the alteration in anatomical structure; cf. Kuster, "Pathologische Pflanzenanatomie", Jena, 1903.) If we look closely into the development of a flowering plant, we notice that in a given species differently formed organs occur in definite positions. In a potato plant colourless runners are formed from the base of the main stem which grow underground and produce tubers at their tips: from a higher level foliage shoots arise nearer the apex. External appearances suggest that both the place of origin and the form of these organs were predetermined in the egg-cell or in the tuber. But it was shown experimentally by the well-known investigator Knight (Knight, "Selection from the Physiological and Horticultural Papers", London, 1841.) that tubers may be developed on the aerial stem in place of foliage shoots. These observations were considerably extended by Vochting. (Vochting, "Ueber die Bildung der Knollen", Cassel, 1887; see also "Bot. Zeit." 1902, 87.) In one kind of potato, germinating tubers were induced to form foliage shoots under the influence of a higher temperature; at a lower temperature they formed tuber-bearing shoots. Many other examples of the conversion of foliage-shoots into runners and rhizomes, or vice versa, have been described by Goebel and others. As in the asexual reproduction of algae quantitative alteration in the amount of moisture, light, temperature, etc. determines whether this or that form of shoot is produced. If the primordia of these organs are exposed to altered conditions of nutrition at a sufficiently early stage a complete substitution of one organ for another is effected. If the rudiment has reached a certain stage in
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