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ation is by far the most important, and most of the others are in a sense merely special cases of this fundamental law. To this law of differentiation is due the increase in complexity or perfection of organisation which is shown by all the animal series. Bronn himself recognised the great similarity of this law of progressive differentiation to Milne-Edwards' principle of the division of labour; he seems, however, to have arrived at it independently. Bronn's third factor in the production of variety of form is adaptation to environment, or better, functional response to environment. Bronn gives an excellent account of adaptational modifications and calls attention, just as Milne-Edwards did, to the numerous analogies of structure which adaptation brings about. He works out the interesting view that there is some connection between classificatory groups and adaptational forms, especially such as are connected with the function of locomotion:--"Based upon a common characteristic method of locomotion are whole or nearly whole sub-phyla (Hexapoda), classes (mammals and reptiles, birds, fishes, gastropods, pteropods, brachiopods, Bryozoa, Rotifera, jelly-fish, polypes, sponges), sub-classes (mobile and immobile lamellibranchs, echinoderms, walking and swimming Crustacea, parasitic and free-living worms, and so on), often, however, only orders and quite small groups (snakes, eels, bats, sepias, medusae, etc.)" (p. 141). It was characteristic of the 'forties and 'fifties that transcendental anatomy, along with Nature-philosophy, went rather out of fashion, its false simplicities and premature generalisations being overwhelmed by the flood of new discoveries. A few stalwarts indeed upheld transcendental views. We have already discussed the morphological system built up by Richard Owen in the late 'forties, a system transcendental in its main lines. We have seen the vertebral theory of the skull still maintained in the 'fifties by such men as Reichert and Koelliker, and we find J. V. Carus in 1853[314] taking it as almost conclusively proved.[315] We may mention, too, as showing clear marks of the influence of transcendental ideas, L. Agassiz's work on the principles of classification.[316] And Serres, who was Geoffroy's chief disciple, recanted not a whit of his doctrine of recapitulation, but re-affirmed and expanded it from time to time, and particularly in a lengthy memoir published in 1860.[317] But in general we may
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