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ches that the green matter originally arises within the primary chlorophyll- or phycochrom-bearing cellule, and consequently is not intruded from any external quarter, nor arises in any way from any parasitism of any kind. The cellule at first is observed to be empty, and then, by the aid of secretion, green matter is gradually produced in the cavity and assumes a definite form. It can, therefore, be very easily and evidently demonstrated that the origin of green matter in lichens is entirely the same as in other plants." On another occasion, and in another place, the same eminent lichenologist remarks,[N] as to the supposed algoid nature of gonidia--"that such an unnatural existence as they would thus pass, enclosed in a prison and deprived of all autonomous liberty, is not at all consonant with the manner of existence of the other algae, and that it has no parallel in nature, for nothing physiologically analogous occurs anywhere else. Krempelhuber has argued that there are no conclusive reasons against the assumption that the lichen-gonidia may be self-developed organs of the lichen proper rather than algae, and that these gonidia can continue to vegetate separately, and so be mistaken for unicellular algae." In this Th. Fries seems substantially to concur. But there is one strong argument, or rather a repetition of an argument already cited, placed in a much stronger light, which is employed by Nylander in the following words:--"So far are what are called algae, according to the turbid hypothesis of Schwendener, from constituting true algae, that on the contrary it may be affirmed that they have a lichenose nature, whence it follows that these pseudo-algae are in a systematic arrangement to be referred rather to the lichens, and that the class of algae hitherto so vaguely limited should be circumscribed by new and truer limits." As to another phase in this question, there are, as Krempelhuber remarks, species of lichens which in many countries do not fructify, and whose propagation can only be carried on by means of the soredia, and the hyphae of such could in themselves alone no more serve for propagation than the hyphae from the pileus or stalk of an Agaric, while it is highly improbable that they could acquire this faculty by interposition of a foreign algal. On the other hand he argues: "It is much more conformable to nature that the gonidia, as self-developed organs of the lichens, should, like the spores, enab
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