ome point of the decomposition. It
rests here for a second or two, and repeats the process; and this is
taking place by what seems almost like rhythmic movement all over the
rotting tissue. The results are scarcely visible in the mass. But if a
group of these organisms be watched, attached to a small particle of
the fermenting tissue, it will be seen to gradually diminish, and at
length to disappear.
Now, there are at least two other similar forms, one of which,
_Heteromita uncinata_, is similar in action, and the other of which,
_Dallingeria drysdali_, is much more powerful, being possessed of a
double anchor, and springing down upon the decadent mass with
relatively far greater power.
Now, it is under the action of these last forms that in a period
varying from one month to two or three the entire substance of the
organic tissues disappears, and the decomposition has been designated
by me "exhausted"; nothing being left in the vessel but slightly
noxious and pale gray water, charged with carbonic acid, and a fine,
buff colored, impalpable sediment at the bottom.
My purpose is not, by this brief notice, to give an exhaustive, or
even a sufficient account, of the progress of fermentative action, by
means of saprophytic organisms, on great masses of tissue; my
observations have been incidental, but they lead me to the conclusion
that the fermentative process is not only not carried through by what
are called saprophytic bacteria, but that a _series_ of fermentative
organisms arise, which succeed each other, the earlier ones preparing
the pabulum or altering the surrounding medium, so as to render it
highly favorable to a succeeding form. On the other hand, the
succeeding form has a special adaptation for carrying on the
fermentative destruction more efficiently from the period at which it
arises, and thus ultimately of setting free the chemical elements
locked up in dead organic compounds.
That these later organisms are saprophytic, although not bacterial,
there can be no doubt. A set of experiments, recorded by me in the
proceedings of this society some years since, would go far to
establish this (_Monthly Microscopical Journal_, 1876, p. 288). But it
may be readily shown, by extremely simple experiments, that these
forms will set up fermentative decomposition rapidly if introduced in
either a desiccated or living condition, or in the spore state, into
suitable but sterilized pabulum.
Thus while we have spec
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