2. No transitional forms of animals are met with among the organic
remains of earlier epochs."
Upon this, Professor Kolliker remarks that the absence of transitional
forms in the fossil world, though not necessarily fatal to Darwin's
views, weakens his case.
"3. The struggle for existence does not take place."
To this objection, urged by Pelzeln, Kolliker, very justly, attaches no
weight.
"4. A tendency of organisms to give rise to useful varieties, and a
natural selection, do not exist.
"The varieties which are found arise in consequence of manifold external
influences, and it is not obvious why they all, or partially, should be
particularly useful. Each animal suffices for its own ends, is perfect
of its kind, and needs no further development. Should, however, a
variety be useful and even maintain itself, there is no obvious
reason why it should change any further. The whole conception of the
imperfection of organisms and the necessity of their becoming perfected
is plainly the weakest side of Darwin's Theory, and a pis aller
(Nothbehelf) because Darwin could think of no other principle by which
to explain the metamorphoses which, as I also believe, have occurred."
Here again we must venture to dissent completely from Professor
Kolliker's conception of Mr. Darwin's hypothesis. It appears to us to be
one of the many peculiar merits of that hypothesis that it involves no
belief in a necessary and continual progress of organisms.
Again, Mr. Darwin, if we read him aright, assumes no special tendency of
organisms to give rise to useful varieties, and knows nothing of
needs of development, or necessity of perfection. What he says is, in
substance: All organisms vary. It is in the highest degree improbable
that any given variety should have exactly the same relations to
surrounding conditions as the parent stock. In that case it is either
better fitted (when the variation may be called useful), or worse
fitted, to cope with them. If better, it will tend to supplant the
parent stock; if worse, it will tend to be extinguished by the parent
stock.
If (as is hardly conceivable) the new variety is so perfectly adapted
to the conditions that no improvement upon it is possible,--it will
persist, because, though it does not cease to vary, the varieties will
be inferior to itself.
If, as is more probable, the new variety is by no means perfectly
adapted to its conditions, but only fairly well adapted to them, it w
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