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of the receptive organs with the efferent nerves that it is not improbable that, through the central nervous organ, each receptive organ is connected with every motor nerve of the whole nervous system,--the facts of strychnine poisoning show that if this is not literally true it is at least approximately so. Hence one of the goals to which each afferent fibre from a receptive organ leads is a number of motor nerves. Their conducting paths must, therefore, converge in passing to the starting-points of the motor nerves; because these latter are instruments common to the use of a number of different receptive organs in so far as they excite reflex actions. On the other hand those of their conducting paths which are concerned in the genesis of sensation, instead of converging, diverge, at least as far as the _cortex cerebri_, or if not divergent, remain separate. These considerations would make it appear likely that the conducting path from each receptive organ divides in the central nervous system into two main lines, one of which goes off to its own particular region of the _cortex cerebri_ whither run conductors only of similar sensual species to itself, while the other main line passes with many others to a great motor station where, as at a telephone exchange, coordinate use of the outgoing lines is assured to them all. Now there is in fact a portion of the cortex in mammals the functions of which are so pre-eminently motor, as judged by our present methods, that it is commonly designated the _motor cortex_ (see fig. 24). This region of the cortex occupies in the Primates, including Man, the pre-central gyrus. Among the items of evidence which reveal its motor capabilities are the following. [Illustration: FIG. 24.--Diagram of the Topography of the Main Groups of Foci in the Motor Field of Chimpanzee.] _The Precentral or Motor Region of the Cortex._--The application to it of electric currents excites movements in the skeletal muscles. The movements occur in the half of the body of the side crossed from that of the hemisphere excited. The "motor representation," as it is termed, is in the cortex better described as a representation of definite actions than of particular muscles. The actions "represented" in the top part of the gyrus, namely next the great longitudinal fissure, move the leg; those in the lowest part of the gyrus belong to the tongue and mouth. The topical distribution along the length of the gyrus may
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