pino, however (see Hildebrand's
version, loc. cit.), describes a similar opening of the cup produced by
pressure on the hairs in some genera of the order.)

Down, June 7th [1860].

Best and most beloved of men, I supplicate and entreat you to observe
one point for me. Remember that the Goodeniaceae have weighed like an
incubus for years on my soul. It relates to Scaevola microcarpa. I
find that in bud the indusium collects all the pollen splendidly, but,
differently from Leschenaultia, cannot be afterwards easily opened.
Further, I find that at an early stage, when the flower first opens, a
boat-shaped stigma lies at the bottom of the indusium, and further
that this stigma, after the flower has some time expanded, grows very
rapidly, when the plant is kept hot, and pushes out of the indusium a
mass of pollen; and at same time two horns project at the corners of the
indusium. Now the appearance of these horns makes me suppose that these
are the stigmatic surfaces. Will you look to this? for if they be by the
relative position of the parts (with indusium and stigma bent at right
angles to style) [I am led to think] that an insect entering a flower
could not fail to have [its] whole back (at the period when, as I have
seen, a whole mass of pollen is pushed out) covered with pollen, which
would almost certainly get rubbed on the two horns. Indeed, I doubt
whether, without this aid, pollen would get on to the horns. What
interests me in the case is the analogy in result with the Lobelia, but
by very different means. In Lobelia the stigma, before it is mature,
pushes by its circular brush of hairs the pollen out of the conjoined
anthers; here the indusium collects pollen, and then the growth of the
stigma pushes it out. In the course of about 1 1/2 hour, I found an
indusium with hairs on the outer edge perfectly clogged with pollen, and
horns protruded, which before the 1 1/2 hour had not one grain of pollen
outside the indusium, and no trace of protruding horns. So you will see
how I wish to know whether the horns are the true stigmatic surfaces. I
would try the case experimentally by putting pollen on the horns, but my
greenhouse is so cold, and my plant so small, and in such a little pot,
that I suppose it would not seed...

The little length of stigmatic horns at the moment when pollen is forced
out of the indusium, compared to what they ultimately attain, makes me
fancy that they are not then mature or ready, and if so