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inally assume a single common parent-form (the monophyletic hypothesis), or several (the polyphyletic hypothesis), is wholly immaterial to the essence of the theory of descent; and it is equally immaterial to its fundamental idea what mechanical causes are assumed for the transformation of the varieties. This assumption of a transformation or metamorphosis of species is, however, indispensable, and the theory of descent is very properly called also the "metamorphosis hypothesis," or "doctrine of transmutation;" as well as Lamarckism, after Jean Lamarck, who first founded it in 1809. III. The doctrine of elimination, or the selection theory, as the doctrine especially of "choice of breed or selection," assumes that almost all, or at any rate most, organic species have originated by a process of selection; the artificial varieties under conditions of domestication--as the races of domestic animals and cultivated plants--through artificial choice of breeds; and the natural varieties of animals and plants in their wild state by natural choice of breeds: in the first case, the will of man effects the selection to suit a purpose; in the second, it is effected in a purposeless way by the "struggle for existence." In both cases the transformation of the organic forms takes place through the reciprocal action of the laws of inheritance and of adaptation; in both cases it depends on the survival or selection of the better-qualified minority. This theory of elimination was first clearly recognised and appreciated in its full significance by Charles Darwin in 1859, and the selection-hypothesis which he founded on it is Darwinism properly so called. The relation that these three great theories, which are frequently confounded, bear to one another may, according to the present position of science, be simply defined as follows:--I. Monism, the universal theory of development, or the monistic progenesis-hypothesis, is the one only scientific theory which affords a rational interpretation of the whole universe and satisfies the craving of our human reason for causality, by bringing all natural phenomena into a mechanical causal-connection as parts of a great uniform process of evolution. II. The theory of transmutation, or descent, is an essential and indispensable element in the monistic development hypothesis, because it is the one only scientific theory which rationally explains the origin of organic species--that is to say, by t
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