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e is no final decision between these opposing views. Professor Wilson of Columbia has suggested that both may contain a measure of truth. Again, it is a question whether the finer granules seen within the cell are or are not typical structures, "capable of assimilation, growth, and division, and hence to be regarded as elementary units of structure standing between the cell and the ultimate molecules of living matter." The more philosophical thinkers, like Spencer, Darwin, Haeckel, Michael Foster, August Weismann, and many others, believe that such "intermediate units must exist, whether or not the microscope reveals them to view." Weismann, who has most fully elaborated a hypothetical scheme of the relations of the intracellular units, identifies the larger of these units not with the ordinary granules of the cell, but with a remarkable structure called chromatin, which becomes aggregated within the cell nucleus at the time of cellular division--a structure which divides into definite parts and goes through some most suggestive manoeuvres in the process of cell multiplication. All these are puzzling structures; and there is another minute body within the cell, called the centro-some, that is quite as much so. This structure, discovered by Van Beneden, has been regarded as essential to cell division, yet some recent botanical studies seem to show that sometimes it is altogether wanting in a dividing cell. In a word, the architecture of the cell has been shown by modern researches to be wonderfully complicated, but the accumulating researches are just at a point where much is obscure about many of the observed phenomena. The immediate future seems full of promise of advances upon present understanding of cell processes. But for the moment it remains for us, as for preceding generations, about the most incomprehensible, scientifically speaking, of observed phenomena, that a single microscopic egg cell should contain within its substance all the potentialities of a highly differentiated adult being. The fact that it does contain such potentialities is the most familiar of every-day biological observations, but not even a proximal explanation of the fact is as yet attainable. THE ANCESTRY OF THE MAMMALS Turning from the cell as an individual to the mature organism which the cell composes when aggregated with its fellows, one finds the usual complement of open questions, of greater or less significance, focalizing
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