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g others (nine in all) to the following conclusions: "First, that the unsymmetrical spiral forms of the shells of these and of all the Mollusca probably resulted from the action of the laws of heredity, modified by gravitation. "Second, that there are many characteristics in these shells and in other groups, which are due solely to the uniform action of the physical influence of the immediate surroundings, varying with every change of locality, but constant and uniform within each locality. "Third, that the Darwinian law of Natural Selection does not explain these relations, but applies only to the first stages in the establishment of the differences between forms or species in the same locality. That its office is to fix these in the organization and bring them within the reach of the laws of heredity." These views we find reiterated in his later palaeontological papers. Hyatt's views on acceleration were adopted by Neumayr.[211] Waagen,[212] from his studies on the Jurassic cephalopods, concludes that the factors in the evolution of these forms were changes in external conditions, geographical isolation, competition, and that the fundamental law was not that of Darwin, but "the law of development." Hyatt has also shown that at first evolution was rapid. "The evolution is a purely mechanical problem in which the action of the habitat is the working agent of all the major changes; first acting upon the adult stages, as a rule, and then through heredity upon the earlier stages in successive generations." He also shows that as the primitive forms migrated and occupied new, before barren, areas, where they met with new conditions, the organisms "changed their habits and structures rapidly to accord with these new conditions."[213] While the palaeontological facts afford complete and abundant proofs of the modifying action of changes in the environment, Hyatt, in 1877, from his studies on sponges,[214] shows that the origin of their endless forms "can only be explained by the action of physical surroundings directly working upon the organization and producing by such direct action the modifications or common variations above described." Mr. A. Agassiz remarks that the effect of the nature of the bottom of the sea on sponges and rhizopods "is an all-important factor in modifying the organism."[215] While Hyatt's studies were chiefly on the ammonites, molluscs, and existing sponges, Cope
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