so in the tissue of the stigma, where in
accordance with the compression of the utriculi, it has an intermediate
form, being neither so much flattened as in the epidermis nor so convex
as it is in the internal tissue of the column.
"I may here remark that I am acquainted with one case of apparent
exception to the nucleus being solitary in each utriculus or
cell--namely, in Bletia Tankervilliae. In the utriculi of the stigma of
this plant, I have generally, though not always, found a second areola
apparently on the surface, and composed of much larger granules than the
ordinary nucleus, which is formed of very minute granular matter, and
seems to be deep seated.
"Mr. Bauer has represented the tissue of the stigma, in the species of
Bletia, both before and, as he believes, after impregnation; and in the
latter state the utriculi are marked with from one to three areolae of
similar appearance.
"The nucleus may even be supposed to exist in the pollen of this family.
In the early stages of its formation, at least a minute areola is of
ten visible in the simple grain, and in each of the constituent parts
of cells of the compound grain. But these areolae may perhaps rather be
considered as merely the points of production of the tubes.
"This nucleus of the cell is not confined to orchideae, but is equally
manifest in many other monocotyledonous families; and I have even
found it, hitherto however in very few cases, in the epidermis of
dicotyledonous plants; though in this primary division it may perhaps
be said to exist in the early stages of development of the pollen. Among
monocotyledons, the orders in which it is most remarkable are Liliaceae,
Hemerocallideae, Asphodeleae, Irideae, and Commelineae.
"In some plants belonging to this last-mentioned family, especially
in Tradascantia virginica, and several nearly related species, it is
uncommonly distinct, not in the epidermis and in the jointed hairs of
the filaments, but in the tissue of the stigma, in the cells of the
ovulum even before impregnation, and in all the stages of formation
of the grains of pollen, the evolution of which is so remarkable in
tradascantia.
"The few indications of the presence of this nucleus, or areola, that I
have hitherto met with in the publications of botanists are chiefly in
some figures of epidermis, in the recent works of Meyen and Purkinje,
and in one case, in M. Adolphe Broigniart's memoir on the structure of
leaves. But so lit
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