round; supernumerary tubercles small, in single row on
proximal segments of each digit; toes moderately long and slender;
length of toes from shortest to longest, 1-2-5-3-4, the fifth nearly
as long as third; toes nearly fully webbed; web extending to base of
penultimate phalanx of fourth toe and to discs on other toes; discs
small, about two-thirds size of those on fingers. Anal opening above
middle of thighs; anal sheath short, deeply creased medially; heavy
transverse dermal fold above anus; no large anal tubercles. Skin of
all dorsal surfaces, chin, and ventral surfaces of limbs, except
proximal parts of thighs, smooth; belly and proximal parts of thighs
areolate; thoracic fold present, weak. Tongue elliptical, slightly
longer than wide, not notched behind, and free posteriorly for about
one-fourth of its length; vomerine teeth 3-3, situated on small,
widely separated, transverse ridges between rather small elliptical
inner nares; no vocal slits.

Color (in alcohol) dark brown with irregular darker reticulations on
dorsal surfaces of head, body, and limbs; flanks brown with small
creamy white spots; posterior surfaces of thighs dark brown; chin
creamy tan; belly grayish brown with cream-colored flecks; ventral
surfaces of limbs pale brown; webbing on feet gray; small white spots
in anal region.

Color (in life) chocolate brown with darker brown reticulations and
irregular blotches above; flanks brown with yellow spots; belly gray
to grayish brown with faint cream-colored spots; iris a deep bronze
color.

_Variation._--In males the total number of vomerine teeth varies from
4 to 7. In many specimens the vomerine ridges are larger and more
closely approximated medially than in the specimen described above.
Females attain snout-vent lengths of 51 mm., have as many as 9
vomerine teeth, and have a proportionately larger tympanum than males.
Some of the largest specimens of both sexes have indistinct
cream-colored pustules scattered on the ventral surface of the
forearm. Some individuals have nearly uniform grayish brown ventral
surfaces; in others the chin, as well as the abdomen, is brown with
cream-colored spots. The dorsal surfaces of some specimens are nearly
uniform dark brown with no reticulations. In others the dorsum is
paler brown with distinct darker mottling; in some of these there is
little mottling laterally, so that there is the effect of an
irregular, pale brown, dorsolateral stripe.

_Tadpoles._--Th