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ger ones the muscles are beginning to assume this function and the animal moves by writhing. The bilateral symmetry has arisen in connection with this mode of locomotion and is thus a mark of important progress. In the turbellaria we find for the first time a true body-wall distinct from underlying organs. The outer layer of this is a ciliated epithelium or layer of cells. Under this an elastic membrane may occur. Then come true body muscles, running transversely, longitudinally and dorso-ventrally. Between the external transverse and the internal longitudinal layers we often find two muscular layers whose fibres run diagonally. The body is well provided with muscles, but their arrangement is still far from economical or effective. Within the body-wall is the parenchym. This is a spongy mass of connectile tissue in which the other organs are embedded. The mouth lies in the middle, or near the front of the ventral surface. The intestine varies in form, but is provided with its own layers of longitudinal and transverse muscles, and usually has paired pouches extending out from it into the body parenchym. These seem to distribute the dissolved nutriment; hence the whole cavity is still often called a gastro-vascular cavity as serving both digestion and circulation. There is no anal opening, but indigestible material is still cast out through the mouth. The animal can gain sufficient oxygen to supply its muscles and nerves, which are the principal seats of combustion, through the external surface. It has, therefore, no special respiratory organs. But the waste matter of the muscles cannot escape so easily, for these are becoming deeper seated. Hence we find an excretory system consisting of two tubes with many branches in the parenchym, and discharging at the rear end of the body. This again is a sign that the muscles are becoming more important, for the excretory system is needed mainly to remove their waste. These tubes maybe only greatly enlarged glands of the skin. [Illustration: 5. TURBELLARIAN. LANG. _va_ and _ha_, front and rear branches of gastro-vascular cavity; _ph_, pharynx. The dark oval with fine branches represents the nervous system.] The nervous system consists of a plexus of fibres and cells, the cells originating impulses and the fibres conveying them. But this much was present in hydra also. Here the front end of the body goes foremost and is continually coming in contact with new con
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