ched the pterygoid, and either met the latter or had
a ligamentous connection with it. As nearly as can be determined, the
total length of the maxillary is approximately 12.0 mm.
The teeth on the maxillary are small and seem to be in two longitudinal
rows. The palatine bears two large, grooved teeth anteriorly; the first
is approximately 1.0 mm. posteromedial to the naris and the second is
about 3.0 mm. posterior and slightly lateral to the naris. The flat
ventral surfaces of the palatine and pterygoid bear numerous small
teeth distributed as shown in Fig. 4.
_Braincase and Occipital Region_ (Fig. 5)
The parts of the neurocranium are scattered, disconnected and
incomplete, but it is possible to make out a number of features of the
otico-occipital section with fair assurance. In posterior view the
notochordal canal and foramen magnum are confluent with each other, and
of great size relative to the skull as a whole. The notochordal canal
measures 2.8 mm. in diameter, and the foramen magnum about 4.0 mm. The
crescent-shaped supraoccipital rests on the upright ends of the
exoccipitals, but between the latter and the basioccipital no sutures
can be seen. Probably the whole posterior surface of the braincase
slanted posteroventrally; consequently the rim of the notochordal canal
was about 3.0 mm. behind the margin of the parietals.
The U-shaped border of the notochordal canal is a thick, rounded bone,
comparable in appearance to the U-shaped intercentra of the vertebrae.
This bone apparently rested upon a thinner, troughlike piece (Fig. 5 B)
forming the floor of the braincase. The latter is broad, shallow,
concave, open midventrally and narrowing anteriorly to form a pair of
articular processes. Since no sutures can be seen in this structure, it
probably is the ventral, ossified portion of the basioccipital. Watson
(1926, Fig. 4 B) illustrates the floor of the braincase in
_Eusthenopteron_, with its more lateral, anterior portion labelled
prootic, but in our specimen the corresponding part could scarcely have
formed the anterior wall of the otic capsule, being entirely in the
plane of the floor. The two articular surfaces anteriorly near the
midline suggest that a movable joint existed between the
otico-occipital part of the braincase and the ethmosphenoid part, as in
Rhipidistia (Romer, 1937). We have found nothing in the specimen that
could be referred to the ethmosphenoid; it may have been unossified.
[Illustrati
|