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rganism taken conjointly with the changes of geology, which are steadily (and sometimes suddenly, as when an isthmus at last separates) in progress, that occasionally organisms must suddenly be introduced into new regions, where, if the conditions of existence are not so foreign as to cause its extermination, it will often be propagated under circumstances still more closely analogous to those of domestication; and therefore we expect will evince a tendency to vary. It appears to me quite _inexplicable_ if this has never happened; but it can happen very rarely. Let us then suppose that an organism by some chance (which might be hardly repeated in 1000 years) arrives at a modern volcanic island in process of formation and not fully stocked with the most appropriate organisms; the new organism might readily gain a footing, although the external conditions were considerably different from its native ones. The effect of this we might expect would influence in some small degree the size, colour, nature of covering &c., and from inexplicable influences even special parts and organs of the body. But we might further (and <this> is far more important) expect that the reproductive system would be affected, as under domesticity, and the structure of the offspring rendered in some degree plastic. Hence almost every part of the body would tend to vary from the typical form in slight degrees, and in no determinate way, and therefore _without selection_ the free crossing of these small variations (together with the tendency to reversion to the original form) would constantly be counteracting this unsettling effect of the extraneous conditions on the reproductive system. Such, I conceive, would be the unimportant result without selection. And here I must observe that the foregoing remarks are equally applicable to that small and admitted amount of variation which has been observed in some organisms in a state of nature; as well as to the above hypothetical variation consequent on changes of condition. Let us now suppose a Being{224} with penetration sufficient to perceive differences in the outer and innermost organization quite imperceptible to man, and with forethought extending over future centuries to watch with unerring care and select for any object the offspring of an organism produced under the foregoing circumstances; I can see no conceivable reason why he could not form a new race (or several were he to separate the stock o
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