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of development and a group of hypothetical germinal determiners that tend to be associated within the germ. The presence or absence of a determiner in a germ is thus the primary cause of the corresponding presence or absence of a certain characteristic in the adult organism. But whatever the essential nature of the characteristic in this respect, whether simple or complex, we know further that every organismal characteristic is subject to variation. In any group of human individuals, for example, we can find persons of different stature, different weight, with fingers of different length and form, with heads of different size and shape, hair and eyes of different shades, different blood pressures, pulse rates, digestive possibilities, different degrees of determination, cheerfulness, alertness, and so forth. This fact of variation is not limited to the comparison of the individuals of a given group or generation among themselves, but successive generations considered as the units of comparison show the same sort of thing. And further successive broods from the same parents exhibit this same phenomenon of variation when compared with one another. Variation is a universal fact--not only among organic things but in the inorganic world as well. The variation which any company of persons shows in stature is paralleled by the variation in the diameter of the grains in a handful of sand, or of the drops in a rainstorm. When we examine the phenomena of variation carefully we find that they are of two quite distinct categories. The first kind of variation, that which we most frequently think of as "variation," should properly be termed _variability_. Differences of this type are small _fluctuations_ in any and every character, centering about an average or mean, which is itself fairly definite and fixed--less subject to variation in different groups or through successive generations. For example, if we measure by inches the stature of a thousand or more persons chosen at random we find that they may vary from fifty-four to seventy-six inches; the most frequent heights might be about sixty-nine and sixty-four inches among the men and women respectively. The results of such a measurement may be expressed graphically as in Figure 3, which is an expression of the measurement of 1,052 mothers. The measurement of almost any characteristic in a large group of any organisms usually gives a result of the kind figured. The most s
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