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ssive of the twilight of the forest. The skin of the female is about the shade of that of the Southern European of to-day; that of the male is darker. The most interesting of the three figures is the young ape-mother, who reclines against a tree-trunk and offers her breast to her first-born. The expression of the face is remarkable; happiness at the possession of the child mingles with misgiving for its future. The tear which trembles upon her cheek seems indicative of the flood of tears which is to run down the history of her descendants. The father has less of this feeling, and stands upright beside his wife and child and looks down upon them with an air of pride and paternal joy. The original painting is owned by the celebrated Darwinian philosopher Ernest Haeckel of Jena.] But at that time I overlooked one problem of great importance; and it is astonishing to me, except on the principle of Columbus and his egg, how I could have overlooked it and its solution. This problem is the tendency in organic beings descended from the same stock to diverge in character as they become modified. That they have diverged greatly is obvious from the manner in which species of all kinds can be classed under genera, genera under families, families under sub-orders, and so forth: and I can remember the very spot in the road, whilst in my carriage, when to my joy the solution occurred to me; and this was long after I had come to Down. The solution, as I believe, is that the modified offspring of all dominant and increasing forms tend to become adapted to many and highly diversified places in the economy of nature. Early in 1856 Lyell advised me to write out my views pretty fully, and I began at once to do so on a scale three or four times as extensive as that which was afterwards followed in my 'Origin of Species'; yet it was only an abstract of the materials which I had collected, and I got through about half the work on this scale. But my plans were overthrown, for early in the summer of 1858 Mr. Wallace, who was then in the Malay Archipelago, sent me an essay 'On the Tendency of Varieties to depart Indefinitely from the Original Type'; and this essay contained exactly the same theory as mine. Mr. Wallace expressed the wish that if I thought well of his essay, I should send it to Lyell for perusal. The circumstances under which I consented, at the request of Lyell and Hooker, to allow of an abstract from my MS., together with a l
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