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erations arising before the next upheaval, the sensible or insensible alterations thus produced in each species would become organised--there would be a more or less complete adaptation to the new conditions. The next upheaval would superinduce further organic changes, implying wider divergences from the primary forms; and so repeatedly. But now let it be observed that the revolution thus resulting would not be a substitution of a thousand more or less modified species for the thousand original species; but in place of the thousand original species there would arise several thousand species, or varieties, or changed forms. Each species being distributed over an area of some extent, and tending continually to colonise the new area exposed, its different members would be subject to different sets of changes. Plants and animals spreading towards the equator would not be affected in the same way with others spreading from it. Those spreading towards the new shores would undergo changes unlike the changes undergone by those spreading into the mountains. Thus, each original race of organisms, would become the root from which diverged several races differing more or less from it and from each other; and while some of these might subsequently disappear, probably more than one would survive in the next geologic period: the very dispersion itself increasing the chances of survival. Not only would there be certain modifications thus caused by change of physical conditions and food, but also in some cases other modifications caused by change of habit. The fauna of each island, peopling, step by step, the newly-raised tracts, would eventually come in contact with the faunas of other islands; and some members of these other faunas would be unlike any creatures before seen. Herbivores meeting with new beasts of prey, would, in some cases, be led into modes of defence or escape differing from those previously used; and simultaneously the beasts of prey would modify their modes of pursuit and attack. We know that when circumstances demand it, such changes of habit _do_ take place in animals; and we know that if the new habits become the dominant ones, they must eventually in some degree alter the organisation. Observe, now, however, a further consequence. There must arise not simply a tendency towards the differentiation of each race of organisms into several races; but also a tendency to the occasional production of a somewhat high
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