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ng the Middle and Upper Eocene epochs. Hence it becomes highly probable that the well-known similarities, and no less remarkable differences between the present Faunae of India and South Africa have arisen in some such fashion as the following. Some time during the Miocene epoch, possibly when the Himalayan chain was elevated, the bottom of the nummulitic sea was upheaved and converted into dry land, in the direction of a line extending from Abyssinia to the mouth of the Ganges. By this means, the Dekhan on the one hand, and South Africa on the other, became connected with the Miocene dry land and with one another. The Miocene mammals spread gradually over this intermediate dry land; and if the condition of its eastern and western ends offered as wide contrasts as the valleys of the Ganges and Arabia do now, many forms which made their way into Africa must have been different from those which reached the Dekhan, while others might pass into both these sub-provinces. That there was a continuity of dry land between Europe and North America during the Miocene epoch, appears to me to be a necessary consequence of the fact that many genera of terrestrial mammals, such as _Castor, Hystrix, Elephas, Mastodon, Equus, Hipparion, Anchitherium, Rhinoceros, Cervus, Amphicyon, Hyoenarctos_, and _Machairodus_, are common to the Miocene formations of the two areas, and have as yet been found (except perhaps _Anchitherium_) in no deposit of earlier age. Whether this connection took place by the east, or by the west, or by both sides of the Old World, there is at present no certain evidence, and the question is immaterial to the present argument; but, as there are good grounds for the belief that the Australian province and the Indian and South-African sub-provinces were separated by sea from the rest of Arctogaea before the Miocene epoch, so it has been rendered no less probable, by the investigations of Mr. Carrick Moore and Professor Duncan, that Austro- Columbia was separated by sea from North America during a large part of the Miocene epoch. It is unfortunate that we have no knowledge of the Miocene mammalian fauna of the Australian and Austro-Columbian provinces; but, seeing that not a trace of a Platyrrhine Ape, of a Procyonine Carnivore, of a characteristically South-American Rodent, of a Sloth, an Armadillo, or an Ant-eater has yet been found in Miocene deposits of Arctogaea, I cannot doubt that they already existed in the M
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