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eneration to another; the question was, Why do they vary? and do these variations really represent new characters comparable to new species in the making? or are they, so to speak, but an elastic reaction of the internal vital elasticity of the organism, all the while latent and only seeking a favorable expression, to return again under other conditions to the former type? The effort to reduce these variations to law and system was pursued by thousands of investigators, with varying but at all times perplexing and disappointing results. But in the year 1900 the scientific world awoke to the surprising fact that a patient obscure investigator had already solved most of the puzzles of variation and heredity some thirty-five years before. Gregor Mendel, born a peasant boy, trained as a monk, and afterwards appointed Abbot of Bruenn, had in the year 1865 published the results of his experiments in breeding, which had been ignored or forgotten until rediscovered in 1900 by de Vries and two others simultaneously. From this point Mendelism, as it is now called, has steadily gained ground, until at the present time it can be said to be the dominating conception among biologists the world over regarding the problems of heredity. Mendel worked chiefly with peas, crossing different varieties. In his methods of investigation he differed from all previous investigators in concentrating his attention upon a single pair of alternative or contrasted characters at a time, and observing how these alternative characters are transmitted. Thus when he crossed a tall with a dwarf, giving attention to this pair of contrasted characters alone, he found that all the first hybrid generation were talls, with no dwarfs and no intermediates. Accordingly he called the tall character _dominant_, and the dwarf character _recessive_, and a pair of contrasted characters which act in this way are now called _factors_ or sometimes called _unit characters_. But on allowing these hybrids to cross-fertilize one another in the usual way, Mendel found that in the second generation of hybrids there were _always_ _three talls to one dwarf_ out of every four. Further experiments proved that these dwarfs of the second hybrid generation _always bred true_, that is, one out of four; and that one out of the remaining talls always bred true, making another quarter of the total; while the remaining fifty per cent. proved to be mixed tails, always acting as did th
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