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orthern land, connecting Europe and America, for their development. We saw, however, that this northern region was singularly warm until long after the spread of the mammals. Other experts, impressed by the parallel development of the mammals and the flowering plants, look to the elevated parts of eastern North America. Such evidence as there is seems rather to suggest that South Africa was the cradle of the placental mammals. We shall find that many of our mammals originated in Africa; there, too, is found to-day the most primitive representative of the Tertiary mammals, the hyrax; and there we find in especial abundance the remains of the mammal-like reptiles (Theromorphs) which are regarded as their progenitors. Further search in the unexplored geological treasures and dense forests of Africa is needed. We may provisionally conceive the placental mammals as a group of the South African early mammals which developed a fortunate variation in womb-structure during the severe conditions of the early Mesozoic. In this new structure they would have no preponderant advantage as long as the genial Jurassic age favoured the great reptiles, and they may have remained as small and insignificant as the Marsupials. But with the fresh upheaval and climatic disturbance at the end of the Jurassic, and during the Cretaceous, they spread northward, and replaced the dying reptiles, as the Angiosperms replaced the dying cycads. When they met the spread of the Angiosperm vegetation they would receive another great stimulus to development. They appear in Europe and North America in the earliest Cretaceous. The rise of the land had connected many hitherto isolated regions, and they seem to have poured over every bridge into all parts of the four continents. The obscurity of their origin is richly compensated by their intense evolutionary interest from the moment they enter the geological record. We have seen this in the case of every important group of plants and animals, and can easily understand it. The ancestral group was small and local; the descendants are widely spread. While, therefore, we discover remains of the later phases of development in our casual cuttings and quarries, the ancestral tomb may remain for ages in some unexplored province of the geological world. If this region is, as we suspect, in Africa, our failure to discover it as yet is all the more intelligible. But these mammals of the early Tertiary are still of
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