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have been much reduced by the general disturbance of conditions, the record is poor. Molluscs and Brachiopods and small fishes fill the list, but are of little instructiveness for us, except that they show a general advance of species. Among the Cephalopods, it is true, we find a notable arrival. On the one hand, a single small straight-shelled Cephalopod lingers for a time with the ancestral form; on the other hand, a new and formidable competitor appears among the coiled-shell Cephalopods. It is the first appearance of the famous Ammonite, but we may defer the description of it until we come to the great age of Ammonites. Of the insects and their fortunes in the great famine we have no direct knowledge; no insect remains have yet been found in Permian rocks. We shall, however, find them much advanced in the next period, and must conclude that the selection acted very effectively among their thousand Carboniferous species. The most interesting outcome of the new conditions is the rise and spread of the reptiles. No other sign of the times indicates so clearly the dawn of a new era as the appearance of these primitive, clumsy reptiles, which now begin to oust the Amphibia. The long reign of aquatic life is over; the ensign of progress passes to the land animals. The half-terrestrial, half-aquatic Amphibian deserts the water entirely (in one or more of its branches), and a new and fateful dynasty is founded. Although many of the reptiles will return to the water, when the land sinks once more, the type of the terrestrial quadruped is now fully evolved, and from its early reptilian form will emerge the lords of the air and the lords of the land, the birds and the mammals. To the uninformed it may seem that no very great advance is made when the reptile is evolved from the Amphibian. In reality the change implies a profound modification of the frame and life of the vertebrate. Partly, we may suppose, on account of the purification of the air, partly on account of the decrease in water surface, the gills are now entirely discarded. The young reptile loses them during its embryonic life--as man and all the mammals and birds do to-day--and issues from the egg a purely lung-breathing creature. A richer blood now courses through the arteries, nourishing the brain and nerves as well as the muscles. The superfluous tissue of the gill-structures is used in the improvement of the ear and mouth-parts; a process that had begun
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