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hanges, and diversification of form and structure, in accordance with changed conditions, their essential nature has probably remained the same from the very dawn of life on the earth. The low aquatic diatomaceae and confervae, together with the lowest fungi and lichens, occupy a similar position in the vegetable kingdom, filling places in nature which would be left vacant if only highly organised plants existed. There is, therefore, no motive power to destroy or seriously to modify them; and they have thus probably persisted, under slightly varying forms, through all geological time. _Extinction of Lower Types among the Higher Animals._ So soon; however, as we approach the higher and more fully developed groups, we see indications of the often repeated extinction of lower by higher forms. This is shown by the great gaps that separate the mammalia, birds, reptiles, and fishes from each other; while the lowest forms of each are always few in number and confined to limited areas. Such are the lowest mammals--the echidna and ornithorhynchus of Australia; the lowest birds--the apteryx of New Zealand and the cassowaries of the New Guinea region; while the lowest fish--the amphioxus or lancelet, is completely isolated, and has apparently survived only by its habit of burrowing in the sand. The great distinctness of the carnivora, ruminants, rodents, whales, bats, and other orders of mammalia; of the accipitres, pigeons, and parrots, among birds; and of the beetles, bees, flies, and moths, among insects, all indicate an enormous amount of extinction among the comparatively low forms by which, on any theory of evolution, these higher and more specialised groups must have been preceded. _Circumstances favourable to the Origin of New Species by Natural Selection._ We have already seen that, when there is no change in the physical or organic conditions of a country, the effect of natural selection is to keep all the species inhabiting it in a state of perfect health and full development, and to preserve the balance that already exists between the different groups of organisms. But, whenever the physical or organic conditions change, to however small an extent, some corresponding change will be produced in the flora and fauna, since, considering the severe struggle for existence and the complex relations of the various organisms, it is hardly possible that the change should not be beneficial to some species and hurtful
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