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ut they do not fuse at any point. These two head-cavities are the only ones to be seen, in this animal, unless the small evaginations from their walls represent other cavities fused with these. Their walls are thin but distinct, and consist of a single layer of cells. These cells are completely filled with their large, round nuclei, so that the wall has the appearance, under higher magnification than is used in this figure, of a band of closely strung, round beads. Figure 4B represents the eighteenth section caudad to the one just described. It passes through that region of the enteron, _ph_, which may be called the pre-oral gut, since it lies cephalad to the now open mouth. Owing to the plane of the section the upper angle of the first gill cleft, _g^1_, is seen on the left, although this would not naturally have been expected in a section through the pre-oral gut. The evagination to form the hypophysis, _p_, is seen against the floor of the forebrain, _fb_. The wall of this region of the enteron is comparatively thin, and consists of not more than two layers of compactly arranged cells with round nuclei. Figure 4C is about forty sections caudad to the one just described. It passes through the mouth, seen as a vertical opening between the two mandibular arches, _md_. The hyomandibular cleft, _g^1_, the only one which opens to the exterior in this embryo, is very wide, and may be traced through a number of sections; in this section the opening is seen only on the left. The pharynx, _ph_, is very wide; as it is followed caudad its ventral opening is gradually closed by the approach of the two mandibular folds. The dorsal wall of this region of the pharynx is very thin, consisting of a single layer of flat cells with round nuclei; while the ventral wall, leading through the mouth and lining the mandibular folds, is composed of two or three layers of compactly arranged cells. Figure 4D is through a plane sixteen sections caudad to the last. In this region, which is just caudad to the otic vesicles, the pharynx has still its rectangular outline, and its walls are of the same character as in the preceding figure. The posterior edges of the hyomandibular clefts are seen projecting in a ventro-lateral direction, _g^1_; while dorsal to these are the wider, second pair of clefts, _g^2_. Where the mandibular folds come together posterior to the mouth, they fuse first at their outer or ventral border, which leaves a deep, na
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