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on swamp out any progressive tendency. Whenever a genius appeared, instead of finding a corresponding genius with which to pair, it mated with the average of its own species. Hence its offspring were nearer the average than it was, and their offspring still nearer. Thus whatever advantage the genius originally possessed gradually sank into the common level. It was Moritz Wagner, a German naturalist, who first insisted that if favorable variations were to amount to anything these possessors must not only mate with others of their same kind, but must also be prevented from mating with the old average group. Accordingly, the belief arose that, under ordinary circumstances, variations returned to the common level. Wherever a varying group became separated by any barrier from mating with the rest of its species, and had only its own kind to pair with, a new species sprang up. This barrier might be a desert, or an impassable mountain range, an arm of the sea, or anything else that the animal could not, or would not, cross. Isolated in this way, the little group that had an advantage in a different direction could develop its tendencies, and a new species would be made of what had been previously only a geographical race. In this matter of geographical isolation Wagner is very strongly supported by the American zooelogist, David Starr Jordan, who believes that no two closely related species of animals ever occupied the same geographical area. Both Wagner and Jordan are ardent admirers of Darwin and his theory of natural selection, but both believe that it is necessary to add the idea of isolation in order to make natural selection effective. George John Romanes, a British naturalist, has added to Wagner's idea of isolation, the expanded conception that there may be isolations that are not geographical. For this phase, Romanes has coined the term physiological isolation. Something in the structure or habit of the animals with the new variation prevents them from mating with the older type. Occasionally it is a difference in the structure of the reproductive organs themselves. This, however, is not the only possible divergence. The mating season in one group may come earlier than that of the other, or may come during the day, while the main group is in the habit of mating at night. Anything which keeps some members of a species separate in their mating from the rest, will result in the course of a longer or shorter time, say
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