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earlier to F.W.A. Miquel. Remarking that "sucking insects (Haustellata)... perform in nature the important duty of maintaining the existence of the vegetable kingdom, at least as far as the higher orders are concerned," he points our that "the appearance in great numbers of haustellate insects occurs at and after the Cretaceous epoch, when the plants with pollen and closed carpels (Angiosperms) are found, and acquire little by little the preponderance in the vegetable kingdom." "Archives Neerlandaises", III. (1868). English translation in "Journ. of Bot." 1869, page 101.) Even with this help the abruptness still remains an almost insoluble problem, though a forecast of floral structure is now recognised in some Jurassic and Lower Cretaceous plants. But the gap between this and the structural complexity and diversity of angiosperms is enormous. Darwin thought that the evolution might have been accomplished during a period of prolonged isolation. Writing to Hooker (1881) he says: "Nothing is more extraordinary in the history of the Vegetable Kingdom, as it seems to me, than the APPARENTLY very sudden or abrupt development of the higher plants. I have sometimes speculated whether there did not exist somewhere during long ages an extremely isolated continent, perhaps near the South Pole." ("Life and Letters", III. page 248.) The present trend of evidence is, however, all in favour of a northern origin for flowering plants, and we can only appeal to the imperfection of the geological record as a last resource to extricate us from the difficulty of tracing the process. But Darwin's instinct that at some time or other the southern hemisphere had played an important part in the evolution of the vegetable kingdom did not mislead him. Nothing probably would have given him greater satisfaction than the masterly summary in which Seward has brought together the evidence for the origin of the Glossopteris flora in Gondwana land. "A vast continental area, of which remnants are preserved in Australia, South Africa and South America... A tract of enormous extent occupying an area, part of which has since given place to a southern ocean, while detached masses persist as portions of more modern continents, which have enabled us to read in their fossil plants and ice-scratched boulders the records of a lost continent, in which the Mesozoic vegetation of the northern continent had its birth." ("Encycl. Brit." (10th edition 1902), Vol.
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