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eir being shut up and deprived of their liberty, or not, we do not know, but they certainly will not breed. What an astounding thing this is, to find one of the most important of all functions annihilated by mere imprisonment! So, again, there are cases known of animals which have been thought by naturalists to be undoubted species, which have yielded perfectly fertile hybrids; while there are other species which present what everybody believes to be varieties [1] which are more or less infertile with one another. There are other cases which are truly extraordinary; there is one, for example, which has been carefully examined,--of two kinds of sea-weed, of which the male element of the one, which we may call A, fertilizes the female element of the other, B; while the male element of B will not fertilize the female element of A; so that, while the former experiment seems to show us that they are 'varieties', the latter leads to the conviction that they are 'species'. When we see how capricious and uncertain this sterility is, how unknown the conditions on which it depends, I say that we have no right to affirm that those conditions will not be better understood by and by, and we have no ground for supposing that we may not be able to experiment so as to obtain that crucial result which I mentioned just now. So that though Mr. Darwin's hypothesis does not completely extricate us from this difficulty at present, we have not the least right to say it will not do so. There is a wide gulf between the thing you cannot explain and the thing that upsets you altogether. There is hardly any hypothesis in this world which has not some fact in connection with it which has not been explained, but that is a very different affair to a fact that entirely opposes your hypothesis; in this case all you can say is, that your hypothesis is in the same position as a good many others. Now, as to the third test, that there are no other causes competent to explain the phenomena, I explained to you that one should be able to say of an hypothesis, that no other known causes than those supposed by it are competent to give rise to the phenomena. Here, I think, Mr. Darwin's view is pretty strong. I really believe that the alternative is either Darwinism or nothing, for I do not know of any rational conception or theory of the organic universe which has any scientific position at all beside Mr. Darwin's. I do not know of any proposition that h
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