ected when they called at night from within crevices of rocks or
from exposed perches on rocks and boulders; some calling frogs were
out of reach on steep rock walls. The call is a single, loud, high
peep.
_Comparisons._--Dixon (1957) recognized six species of
_Tomodactylus_ (_nitidus_, _dilatus_, _albolabris_,
_angustidigitorum_, _fuscus_ and _grandis_) in his revision
of the genus. Another species (_rufescens_) was subsequently
described by Duellman and Dixon (1959). _Tomodactylus
saxatilis_ differs from all the species named immediately
above by the combination of characters given in the
diagnosis. _Tomodactylus saxatilis_ differs from _nitidus_,
_angustidigitorum_ and _grandis_ in having the tips of the
two outer fingers widened and truncate; _saxatilis_ differs
from _dilatus_, _albolabris_, _fuscus_ and _rufescens_ in
having a smooth venter (not pustulate), a contrasting pale
and dark marbled pattern on the back, and a lack of "flash"
colors on the femora.
_Tomodactylus saxatilis_, having lumbar glands, also
resembles three species referred to the genus _Syrrhophus_.
_Tomodactylus macrotympanum_ was described by Taylor
(1940:496, 497) as having a large, moderately distinct lumbar
gland; the species was referred to the genus _Syrrhophus_ by
Dixon (_op. cit._:384). According to Firschein (1954:55),
_Syrrhophus smithi_ and _S. petrophilus_ have elongate lumbar
glands shaped like those in _Tomodactylus_. _Tomodactylus
saxatilis_ resembles _macrotympanum_, _smithi_ and
_petrophilus_ more than it does other species; all four
attain large maximal sizes, and have lumbar glands, mostly
smooth ventral surfaces, three palmar tubercles (sometimes
absent in _saxatilis_), and usually contrasting dorsal
patterns (reduced to flecks and spots in all species except
_saxatilis_). _Tomodactylus saxatilis_ differs from
_macrotympanum_ in having an extensive marbled dorsal pattern
and a small tympanum, and differs from _smithi_ and
_petrophilus_ in having a marbled dorsal pattern and the tips
of the outer two fingers widened and truncate. _Tomodactylus
saxatilis_ differs from all other named species of
_Syrrhophus_ in having conspicuous lumbar glands and in
lacking inguinal glands.
_Remarks._--The characteristics delimiting the genera _Tomodactylus_
and _Syrrhophus_ are not agreed upon
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